; patch choice model from optimal foraging theory ; by Michael Barton, Arizona State University breed [foragers forager] breed [animals animal] foragers-own [energy diet-breadth] animals-own [species food-value processing-costs rank] patches-own [ptimer] globals [rank-list prey-list diversity] to Setup clear-all Setup_Animals Setup_Foragers Setup_Patches end to Go ask foragers [ Move set energy energy - 1 Forage Calculate-Diversity Check-Death ] ask animals [ Move ;;Reproduce ] ask patches [Patch_Color] Do_Plots tick if not any? foragers [stop] end to Setup_Foragers create-foragers init-foragers [ set shape "hunter" set size 2.5 set color white set energy 100 set prey-list [] ; rolling list of prey species taken ] ask foragers [setxy random-xcor random-ycor] ; place the foragers randomly in the world end to Setup_Animals ; Create 4 animal species with different processing costs, food values, birth rates, and initial population densities let total-density (density1 + density2 + density3 + density4) let number1 round (init-prey * density1 / total-density) let number2 round (init-prey * density2 / total-density) let number3 round (init-prey * density3 / total-density) let number4 round (init-prey * density4 / total-density) set rank-list (list (food-value1 - processing-cost1) (food-value2 - processing-cost2) (food-value3 - processing-cost3) (food-value4 - processing-cost4)) set rank-list sort-by [?1 > ?2] rank-list create-animals number1 [ set species 1 set shape "cow" set size 2 set color brown set food-value food-value1 set processing-costs processing-cost1 set rank position (food-value1 - processing-cost1) rank-list + 1 ] create-animals number2 [ set species 2 set shape "rabbit" set size 1.5 set color grey set food-value food-value2 set processing-costs processing-cost2 set rank position (food-value2 - processing-cost2) rank-list + 1 ] create-animals number3 [ set species 3 set shape "fish" set size 1.5 set color blue set food-value food-value3 set processing-costs processing-cost3 set rank position (food-value3 - processing-cost3) rank-list + 1 ] create-animals number4 [ set species 4 set shape "turtle" set size 1.5 set color lime set food-value food-value4 set processing-costs processing-cost4 set rank position (food-value4 - processing-cost4) rank-list + 1 ] ask animals [setxy random-xcor random-ycor] ; place the animals randomly in the world end to Setup_Patches ask patches [set ptimer 20] end to Move rt random 45 lt random 45 fd 1 end to Forage let prey one-of animals-here ;; seek a random animal if prey != nobody [ ;; did we get one? If so, if (energy >= 85 and [rank] of prey = 1) or ;; see how hungry we are and decide whether to take it (energy < 85 and energy >= 70 and [rank] of prey < 3) or (energy < 70 and energy >= 55 and [rank] of prey < 4) or (energy < 55) [ set [pcolor] of patch-here red set [ptimer] of patch-here 0 set energy energy + [food-value] of prey ;; get energy from eating animal set prey-list fput ([species] of prey) prey-list ; add prey-species to running list of prey taken ] ] while [length prey-list > 10] [set prey-list remove-item 10 prey-list] ; manage running list of prey taken end to Patch_Color ifelse ptimer < 20 [set ptimer ptimer + 1] [if pcolor != black [set pcolor black]] end to Calculate-Diversity set diversity 0 if member? 1 prey-list [set diversity diversity + 1] if member? 2 prey-list [set diversity diversity + 1] if member? 3 prey-list [set diversity diversity + 1] if member? 4 prey-list [set diversity diversity + 1] end to Do_Plots set-current-plot "Prey Taken" set-current-plot-pen "species 1" plot length (filter [ ? = 1] prey-list) set-current-plot-pen "species 2" plot length (filter [ ? = 2] prey-list) set-current-plot-pen "species 3" plot length (filter [ ? = 3] prey-list) set-current-plot-pen "species 4" plot length (filter [ ? = 4] prey-list) set-current-plot "Forager Energy" set-current-plot-pen "fenergy" plot (sum [energy] of foragers) end to Check-Death ask foragers [if energy <= 0 [die]] end ;; makes a movie of the model; stops after 500 cycles ;; and exports movie to a file to make-movie ;; prompt user for movie location user-message "First, save your movie file (choose a name ending with .mov)" let path user-new-file if not is-string? path [ stop ] ;; stop if user canceled ;; run the model setup movie-start path movie-grab-view while [ ticks < 500 ] [ go movie-grab-view ] ;; export the movie movie-close user-message (word "Exported movie to " path) end @#$#@#$#@ GRAPHICS-WINDOW 515 60 1137 629 -1 -1 15.0 1 10 1 1 1 0 1 1 1 0 40 0 35 1 1 1 ticks CC-WINDOW 5 644 1146 739 Command Center 0 BUTTON 5 10 71 43 setup Setup NIL 1 T OBSERVER NIL NIL NIL NIL BUTTON 75 10 138 43 run Go T 1 T OBSERVER NIL NIL NIL NIL SLIDER 5 50 180 83 init-foragers init-foragers 1 20 1 1 1 NIL HORIZONTAL SLIDER 5 150 180 183 processing-cost1 processing-cost1 0 10 1 1 1 NIL HORIZONTAL SLIDER 5 115 180 148 food-value1 food-value1 5 100 25 1 1 NIL HORIZONTAL BUTTON 145 10 208 43 step go NIL 1 T OBSERVER NIL NIL NIL NIL BUTTON 260 10 357 43 save movie make-movie NIL 1 T OBSERVER NIL NIL NIL NIL SLIDER 5 185 180 218 density1 density1 0 100 15 1 1 % HORIZONTAL SLIDER 190 150 365 183 processing-cost2 processing-cost2 0 10 3 1 1 NIL HORIZONTAL SLIDER 5 285 180 318 processing-cost3 processing-cost3 0 10 1 1 1 NIL HORIZONTAL SLIDER 190 185 365 218 density2 density2 0 100 25 1 1 % HORIZONTAL SLIDER 5 320 180 353 density3 density3 0 100 40 1 1 % HORIZONTAL SLIDER 190 115 365 148 food-value2 food-value2 5 100 25 1 1 NIL HORIZONTAL SLIDER 5 250 180 283 food-value3 food-value3 5 100 15 1 1 NIL HORIZONTAL SLIDER 190 285 365 318 processing-cost4 processing-cost4 0 10 3 1 1 NIL HORIZONTAL SLIDER 190 320 365 353 density4 density4 0 100 40 1 1 % HORIZONTAL SLIDER 190 250 365 283 food-value4 food-value4 5 100 15 1 1 NIL HORIZONTAL TEXTBOX 15 95 80 113 Species 1 12 0.0 1 SLIDER 190 50 365 83 init-prey init-prey 0 1000 200 1 1 NIL HORIZONTAL PLOT 5 365 510 510 Prey Taken NIL # /10 cycles 0.0 10.0 0.0 10.0 true true PENS "species 1" 1.0 0 -6459832 true "species 2" 1.0 0 -7500403 true "species 3" 1.0 0 -13345367 true "species 4" 1.0 0 -13840069 true MONITOR 375 115 480 160 Forager Energy sum [energy] of foragers 17 1 11 MONITOR 375 165 480 210 Diet Diversity diversity 17 1 11 MONITOR 375 265 440 310 Species 1 length (filter [ ? = 1] prey-list) 0 1 11 TEXTBOX 200 95 260 113 Species 2 12 0.0 1 TEXTBOX 15 230 75 248 Species 3 12 0.0 1 TEXTBOX 200 230 270 248 Species 4 12 0.0 1 MONITOR 440 265 505 310 Species 2 length (filter [ ? = 2] prey-list) 0 1 11 MONITOR 375 310 440 355 Species 3 length (filter [ ? = 3] prey-list) 0 1 11 MONITOR 440 310 505 355 Species 4 length (filter [ ? = 4] prey-list) 0 1 11 TEXTBOX 400 230 495 256 Species Taken\nOver 10 Cycles 11 0.0 1 PLOT 0 510 445 630 Forager Energy NIL energy 0.0 10.0 0.0 10.0 true false PENS "fenergy" 1.0 0 -2674135 true TEXTBOX 650 30 1040 50 Diet Breadth Model (Optimal Foraging Theory) 16 103.0 1 @#$#@#$#@ OVERVIEW ----------- This is an agent-based simulation of the classic "diet breadth model" of optimal foraging theory (see Foley 1985). SIMULATION OPERATION ------------ SETUP: Initialize the number of foragers and total number of prey. FORAGERS: One or more foragers ( selected by the user) are placed randomly and given 100 energy units (eu's) to start with. Each forager begins to move in random directions; each cell moved costs the forager 1 energy unit. PREY: Prey (total determined by , selected by the user) are placed randomly and move randomly. Up to 4 distinct prey species can be defined. The user selects the relative density of the species, its food value (when consumed by a forager), and the costs to process the species before it can be eaten. Prey are ranked according to their net food value = gross food value - processing costs. FORAGING: When a forager encounters prey, she/he decides whether to take it or continue searching for prey. If she/he is not very hungry (energy >= 85), she/he will only take the 1st ranked prey; if she/he is hungrier (energy 70-85), she/he will take 1st or 2nd order prey; if she/he is even hungrier (energy 55-70, she/he will take prey ranked 1st through 3rd; if she/he is very hungry (energy < 55), she/he will take any prey. On taking any prey, the forager received the net food value. A patch turns red briefly to mark when a prey is taken. MONITORING DIET BREADTH: The species of any prey taken is added to the beginning of a running list of the 10 most recent prey taken; if the length of the list is over 10, the last prey on the list is removed. The number of different prey species in the list is monitored as diet breadth. HOW TO USE IT ------------- Set the options (see above). Press "setup". Then press "run". THINGS TO TRY ---------------- Try changing the density of the 1st ranked species or of other species. What happens to diet diversity? Try changing the food values in each cell or the distance moved by each forger each cycle. The classic diet breadth model considers only one forager. What happens if more than one forager are placed in the simulation? EXTENDING THE MODEL ------------------- Other OFT models could be simulated in this way. CREDITS AND REFERENCES ---------------------- C. Michael Barton, Arizona State University For an overview of OFT models, see Foley, R. (1985). Optimality theory in anthropology. 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