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NetLogo Models Library:
Sample Models/Biology

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Fireflies

[screen shot]

If you download the NetLogo application, this model is included. (You can also run this model in your browser, but we don't recommend it; details here.)

## WHAT IS IT?

This model demonstrates a population of fireflies which synchronize their flashing using only the interactions between the individual fireflies. It is a good example of how a distributed system (i.e. a system with many interacting elements, but no 'leader') can coordinate itself without any central coordinator.

Though most species of firefly are not generally known to synchronize in groups, there are some (for example, Pteroptyx cribellata, Luciola pupilla,and Pteroptyx malaccae) that have been observed to do so in certain settings. This model generalizes two main strategies used by such insects to synchronize with each other (phase delay and phase advance synchronization, as described below), retaining the essentials of the strategies while downplaying biological detail.

## HOW IT WORKS

Each firefly constantly cycles through its own clock, flashing at the beginning of each cycle and then resetting the clock to zero once it has reached the maximum. At the start of each simulation all fireflies begin at a random point in their cycles (though they all have the same cycle lengths) and so flashing will occur erratically through the population. As fireflies perceive other flashes around them they are able to use this information to reset their own clocks to try and synchronize with the other fireflies in their vicinity. Each firefly uses the same set of rules to govern its own clock, and depending on the parameters of the simulation, the population may synchronize more or less effectively.

## HOW TO USE IT

GO: starts and stops the simulation.

SETUP: resets the simulation according to the parameters set by the sliders.

NUMBER: sets the number of fireflies to be created.

CYCLE-LENGTH: sets the length of each firefly's clock before it resets to 0.

FLASHES-TO-RESTART: sets the number of flashes a firefly must see in a single tick before its clock resets.

FLASH-LENGTH: sets the duration, in ticks, of each flash.

STRATEGY: sets the synchronization strategy to be used. One value is phase delay, where upon seeing FLASHES-TO-RESTART flashes a firefly will reset its clock to the FLASH-LENGTH tick (just after a flash would normally occur). This causes the firefly to synchronize with the next flash of the firefly it is responding to. The other value is phase advance, where upon seeing FLASHES-TO-RESTART flashes a firefly will reset its clock to zero. This causes the firefly to flash immediately. Under phase advance, fireflies can only begin to reset their clocks during a window which begins two ticks after they have flashed. This assures that flashes do not get stuck in a short cycle where they persistently reset their clocks and stay lit indefinitely.

SHOW-DARK-FIREFLIES: if switch set to on, non-flashing fireflies are displayed in gray. If switch set to off, non-flashing fireflies are colored black and, thus, invisible.

All settings (except SHOW-DARK-FIREFLIES) must be set before pressing the SETUP button. Changes to the sliders (except SHOW-DARK-FIREFLIES) will have no effect on a simulation in progress.

## THINGS TO NOTICE

Using the default settings (number: 1500, cycle-length: 10, flash-length: 1, number-flashes: 1, flashes-to-reset: 1, strategy: "delay"), notice how local clusters of synchronization begin to form. See if you can figure out where each cluster is represented on the plot. As the simulation proceeds, try to determine which local cluster will eventually "take over" the population. Did this cluster originally have the highest spike on the plot?

In phase advance simulations, why do the plots generally top off before the peaks reach the entire population?

In this model fireflies cannot reset their cycle when they are in the middle of a flash. Why was this restriction imposed?

## THINGS TO TRY

Run the model with the "delay"strategy in the default settings. Change FLASHES-TO-RESET to 2, and run the model with the "advance" strategy (while keeping the other settings steady). Which strategy seems more effective? Why?

Try adjusting FLASHES-TO-RESET between 0, 1 and 2 using both phase delay and phase advance settings. Notice that each setting will give a characteristically different plot, and some of them do not allow for synchronization at all (for example, with the delay strategy, contrast FLASHES-TO-RESET set to 1 as opposed to 2). Why does this control make such a difference in the outcome of the simulation?

Changing the number of fireflies in a simulation affects the density of the population (as does adjusting the size of the world). What effect does this have on a simulation?

## EXTENDING THE MODEL

This model explores only two general strategies for attaining synchrony in such cycle-governed fireflies. Can you find any others? Can you improve the existing strategies (i.e., by speeding them up)?

There are many other possible situations in which distributed agents must synchronize their behavior through the use of simple rules. What if, instead of perceiving only other discrete flashes, an insect could sense where another insect was in its cycle (perhaps by hearing an increasingly loud hum)? What kinds of strategies for synchronization might be useful in such a situation?

If all fireflies had adjustable cycle-lengths (initially set to random intervals) would it then be possible to coordinate both their cycle-lengths and their flashing?

Introduce some blind fireflies. How do the strategies fare with this addition?

## NETLOGO FEATURES

Note the use of agentsets to count the number of nearby fireflies that are flashing:

count turtles in-radius 1 with [color = yellow]

## CREDITS AND REFERENCES

Buck, John. (1988). Synchronous Rhythmic Flashing of Fireflies. The Quarterly Review of Biology, September 1988, 265 - 286.

Carlson, A.D. & Copeland, J. (1985). Flash Communication in Fireflies. The Quarterly Review of Biology, December 1985, 415 - 433.

## HOW TO CITE

If you mention this model in a publication, we ask that you include these citations for the model itself and for the NetLogo software:

* Wilensky, U. (1997). NetLogo Fireflies model. http://ccl.northwestern.edu/netlogo/models/Fireflies. Center for Connected Learning and Computer-Based Modeling, Northwestern University, Evanston, IL.
* Wilensky, U. (1999). NetLogo. http://ccl.northwestern.edu/netlogo/. Center for Connected Learning and Computer-Based Modeling, Northwestern University, Evanston, IL.

## COPYRIGHT AND LICENSE

Copyright 1997 Uri Wilensky.

![CC BY-NC-SA 3.0](http://i.creativecommons.org/l/by-nc-sa/3.0/88x31.png)

This work is licensed under the Creative Commons Attribution-NonCommercial-ShareAlike 3.0 License. To view a copy of this license, visit http://creativecommons.org/licenses/by-nc-sa/3.0/ or send a letter to Creative Commons, 559 Nathan Abbott Way, Stanford, California 94305, USA.

Commercial licenses are also available. To inquire about commercial licenses, please contact Uri Wilensky at uri@northwestern.edu.

This model was created as part of the project: CONNECTED MATHEMATICS: MAKING SENSE OF COMPLEX PHENOMENA THROUGH BUILDING OBJECT-BASED PARALLEL MODELS (OBPML). The project gratefully acknowledges the support of the National Science Foundation (Applications of Advanced Technologies Program) -- grant numbers RED #9552950 and REC #9632612.

This model was converted to NetLogo as part of the projects: PARTICIPATORY SIMULATIONS: NETWORK-BASED DESIGN FOR SYSTEMS LEARNING IN CLASSROOMS and/or INTEGRATED SIMULATION AND MODELING ENVIRONMENT. The project gratefully acknowledges the support of the National Science Foundation (REPP & ROLE programs) -- grant numbers REC #9814682 and REC-0126227. Converted from StarLogoT to NetLogo, 2001.

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